Xinjiang Agricultural Sciences ›› 2024, Vol. 61 ›› Issue (11): 2769-2778.DOI: 10.6048/j.issn.1001-4330.2024.11.018
• Plant Protection • Previous Articles Next Articles
CHEN Guoxiang1(), WEI Yang1, GUO Wenchao2, LI Peixuan3, Adili Shataer1(
)
Received:
2024-05-11
Online:
2024-11-20
Published:
2025-01-08
Correspondence author:
Adili Shataer
Supported by:
陈国祥1(), 魏杨1, 郭文超2, 李佩璇3, 阿地力·沙塔尔1(
)
通讯作者:
阿地力·沙塔尔
作者简介:
陈国祥(1998-),男,新疆乌鲁木齐人,硕士研究生,研究方向为害虫综合治理,(E-mail)411895946@qq.com
基金资助:
CLC Number:
CHEN Guoxiang, WEI Yang, GUO Wenchao, LI Peixuan, Adili Shataer. Spatial distribution patterns and sampling techniques of codling moth in walnut orchards[J]. Xinjiang Agricultural Sciences, 2024, 61(11): 2769-2778.
陈国祥, 魏杨, 郭文超, 李佩璇, 阿地力·沙塔尔. 核桃园苹果蠹蛾的空间分布型与抽样技术分析[J]. 新疆农业科学, 2024, 61(11): 2769-2778.
聚集度指标 Aggregation indexes | 均匀分布 Uniform distribution | 随机分布 Random distribution | 聚集分布 Aggregate distribution |
---|---|---|---|
平均拥挤度m* Mean crowding m* | <1 | =1 | >1 |
聚块性指标m*/m Aggregating index m*/m | <1 | =1 | >1 |
丛生指数I Clumping index I | <0 | =0 | >0 |
扩散系数C Diffusion coefficient C | <1 | =1 | >1 |
久野指数CA CA index | <0 | =0 | >0 |
负二项分布指数K Negative binomial distribution index K | <0 | →∞ | >0 |
Tab.1 Criteria for determining population aggregation index
聚集度指标 Aggregation indexes | 均匀分布 Uniform distribution | 随机分布 Random distribution | 聚集分布 Aggregate distribution |
---|---|---|---|
平均拥挤度m* Mean crowding m* | <1 | =1 | >1 |
聚块性指标m*/m Aggregating index m*/m | <1 | =1 | >1 |
丛生指数I Clumping index I | <0 | =0 | >0 |
扩散系数C Diffusion coefficient C | <1 | =1 | >1 |
久野指数CA CA index | <0 | =0 | >0 |
负二项分布指数K Negative binomial distribution index K | <0 | →∞ | >0 |
样地编号 Plot number | 样本数 Sample number | 均值 Mean value (m) | 方差(S2) Variance | 平均拥 挤度 (m*) Mean crowding | 丛生 指数(I) Clumping index | 聚块性 指标 (m*/m) Patch index | 久野 指数 (CA) Cassie index | 扩散 系数 (C) Dispersion coefficient | 负二项 分布指数 (K) Negative binomial distribution index | 分布型 Distribution pattern |
---|---|---|---|---|---|---|---|---|---|---|
1 | 57 | 3.737 | 49.062 | 7.004 | 3.268 | 1.874 | 0.874 | 4.268 | 1.144 | 聚集型 |
2 | 57 | 5.421 | 154.550 | 12.432 | 7.011 | 2.293 | 1.293 | 8.011 | 0.773 | 聚集型 |
3 | 57 | 9.965 | 805.992 | 28.390 | 18.425 | 2.849 | 1.849 | 19.425 | 0.541 | 聚集型 |
4 | 57 | 9.877 | 574.901 | 23.977 | 14.100 | 2.428 | 1.428 | 15.100 | 0.701 | 聚集型 |
5 | 57 | 9.561 | 357.093 | 18.897 | 9.336 | 1.976 | 0.976 | 10.336 | 1.024 | 聚集型 |
6 | 57 | 8.825 | 283.384 | 16.834 | 8.009 | 1.908 | 0.908 | 9.009 | 1.102 | 聚集型 |
7 | 57 | 22.912 | 2 449.260 | 49.490 | 26.578 | 2.160 | 1.160 | 27.578 | 0.862 | 聚集型 |
8 | 57 | 19.860 | 1 362.894 | 36.917 | 17.058 | 1.859 | 0.859 | 18.058 | 1.164 | 聚集型 |
9 | 57 | 20.579 | 1 775.350 | 42.135 | 21.556 | 2.048 | 1.048 | 22.556 | 0.955 | 聚集型 |
10 | 57 | 24.123 | 2 053.123 | 45.311 | 21.189 | 1.878 | 0.878 | 22.189 | 1.139 | 聚集型 |
11 | 57 | 8.597 | 256.016 | 16.001 | 7.404 | 1.861 | 0.861 | 8.404 | 1.161 | 聚集型 |
12 | 57 | 10.105 | 377.296 | 19.424 | 9.319 | 1.922 | 0.922 | 10.319 | 1.084 | 聚集型 |
13 | 57 | 9.018 | 409.245 | 20.230 | 11.212 | 2.243 | 1.243 | 12.212 | 0.804 | 聚集型 |
14 | 57 | 11.632 | 1 143.907 | 33.822 | 22.190 | 2.908 | 1.908 | 23.190 | 0.524 | 聚集型 |
15 | 57 | 6.719 | 227.672 | 15.089 | 8.370 | 2.246 | 1.246 | 9.370 | 0.803 | 聚集型 |
Tab.2 Various aggregation indexes of adult apple moth in walnut garden
样地编号 Plot number | 样本数 Sample number | 均值 Mean value (m) | 方差(S2) Variance | 平均拥 挤度 (m*) Mean crowding | 丛生 指数(I) Clumping index | 聚块性 指标 (m*/m) Patch index | 久野 指数 (CA) Cassie index | 扩散 系数 (C) Dispersion coefficient | 负二项 分布指数 (K) Negative binomial distribution index | 分布型 Distribution pattern |
---|---|---|---|---|---|---|---|---|---|---|
1 | 57 | 3.737 | 49.062 | 7.004 | 3.268 | 1.874 | 0.874 | 4.268 | 1.144 | 聚集型 |
2 | 57 | 5.421 | 154.550 | 12.432 | 7.011 | 2.293 | 1.293 | 8.011 | 0.773 | 聚集型 |
3 | 57 | 9.965 | 805.992 | 28.390 | 18.425 | 2.849 | 1.849 | 19.425 | 0.541 | 聚集型 |
4 | 57 | 9.877 | 574.901 | 23.977 | 14.100 | 2.428 | 1.428 | 15.100 | 0.701 | 聚集型 |
5 | 57 | 9.561 | 357.093 | 18.897 | 9.336 | 1.976 | 0.976 | 10.336 | 1.024 | 聚集型 |
6 | 57 | 8.825 | 283.384 | 16.834 | 8.009 | 1.908 | 0.908 | 9.009 | 1.102 | 聚集型 |
7 | 57 | 22.912 | 2 449.260 | 49.490 | 26.578 | 2.160 | 1.160 | 27.578 | 0.862 | 聚集型 |
8 | 57 | 19.860 | 1 362.894 | 36.917 | 17.058 | 1.859 | 0.859 | 18.058 | 1.164 | 聚集型 |
9 | 57 | 20.579 | 1 775.350 | 42.135 | 21.556 | 2.048 | 1.048 | 22.556 | 0.955 | 聚集型 |
10 | 57 | 24.123 | 2 053.123 | 45.311 | 21.189 | 1.878 | 0.878 | 22.189 | 1.139 | 聚集型 |
11 | 57 | 8.597 | 256.016 | 16.001 | 7.404 | 1.861 | 0.861 | 8.404 | 1.161 | 聚集型 |
12 | 57 | 10.105 | 377.296 | 19.424 | 9.319 | 1.922 | 0.922 | 10.319 | 1.084 | 聚集型 |
13 | 57 | 9.018 | 409.245 | 20.230 | 11.212 | 2.243 | 1.243 | 12.212 | 0.804 | 聚集型 |
14 | 57 | 11.632 | 1 143.907 | 33.822 | 22.190 | 2.908 | 1.908 | 23.190 | 0.524 | 聚集型 |
15 | 57 | 6.719 | 227.672 | 15.089 | 8.370 | 2.246 | 1.246 | 9.370 | 0.803 | 聚集型 |
样地编号 Plot number | 样本数 Sample number | 均值 Mean value (m) | 方差(S2) Variance | 平均拥 挤度 (m*) Mean crowding | 丛生 指数(I) Clumping index | 聚块性 指标 (m*/m) Patch index | 久野 指数 (CA) Cassie index | 扩散 系数 (C) Dispersion coefficient | 负二项 分布指数 (K) Negative binomial distribution index | 分布型 Distribution pattern |
---|---|---|---|---|---|---|---|---|---|---|
1 | 19 | 1.000 | 1.291 | 1.667 | 0.667 | 1.667 | 0.667 | 1.667 | 1.500 | 聚集型 |
2 | 19 | 1.316 | 1.293 | 1.587 | 0.271 | 1.206 | 0.206 | 1.271 | 4.852 | 聚集型 |
3 | 19 | 1.895 | 1.487 | 2.061 | 0.167 | 1.088 | 0.088 | 1.167 | 11.365 | 聚集型 |
4 | 19 | 2.895 | 2.492 | 4.040 | 1.146 | 1.396 | 0.396 | 2.146 | 2.527 | 聚集型 |
5 | 19 | 1.632 | 1.862 | 2.757 | 1.125 | 1.690 | 0.690 | 2.125 | 1.450 | 聚集型 |
6 | 19 | 2.000 | 1.564 | 2.222 | 0.222 | 1.111 | 0.111 | 1.222 | 8.998 | 聚集型 |
7 | 19 | 3.526 | 3.791 | 6.603 | 3.076 | 1.872 | 0.872 | 4.076 | 1.146 | 聚集型 |
8 | 19 | 6.105 | 5.557 | 10.163 | 4.057 | 1.665 | 0.665 | 5.057 | 1.505 | 聚集型 |
9 | 19 | 5.263 | 4.931 | 8.883 | 3.620 | 1.688 | 0.688 | 4.620 | 1.454 | 聚集型 |
10 | 19 | 3.263 | 2.535 | 4.233 | 0.970 | 1.297 | 0.297 | 1.970 | 3.366 | 聚集型 |
11 | 19 | 4.105 | 2.665 | 4.835 | 0.729 | 1.178 | 0.178 | 1.729 | 5.629 | 聚集型 |
12 | 19 | 5.158 | 3.338 | 6.318 | 1.160 | 1.225 | 0.225 | 2.160 | 4.447 | 聚集型 |
13 | 19 | 2.737 | 2.423 | 3.882 | 1.145 | 1.419 | 0.419 | 2.145 | 2.390 | 聚集型 |
14 | 19 | 3.263 | 3.088 | 5.186 | 1.923 | 1.589 | 0.589 | 2.923 | 1.697 | 聚集型 |
15 | 19 | 2.579 | 1.981 | 3.101 | 0.522 | 1.202 | 0.202 | 1.522 | 4.945 | 聚集型 |
Tab.3 Index of various aggregation degree of larval population of apple moth in walnut garden
样地编号 Plot number | 样本数 Sample number | 均值 Mean value (m) | 方差(S2) Variance | 平均拥 挤度 (m*) Mean crowding | 丛生 指数(I) Clumping index | 聚块性 指标 (m*/m) Patch index | 久野 指数 (CA) Cassie index | 扩散 系数 (C) Dispersion coefficient | 负二项 分布指数 (K) Negative binomial distribution index | 分布型 Distribution pattern |
---|---|---|---|---|---|---|---|---|---|---|
1 | 19 | 1.000 | 1.291 | 1.667 | 0.667 | 1.667 | 0.667 | 1.667 | 1.500 | 聚集型 |
2 | 19 | 1.316 | 1.293 | 1.587 | 0.271 | 1.206 | 0.206 | 1.271 | 4.852 | 聚集型 |
3 | 19 | 1.895 | 1.487 | 2.061 | 0.167 | 1.088 | 0.088 | 1.167 | 11.365 | 聚集型 |
4 | 19 | 2.895 | 2.492 | 4.040 | 1.146 | 1.396 | 0.396 | 2.146 | 2.527 | 聚集型 |
5 | 19 | 1.632 | 1.862 | 2.757 | 1.125 | 1.690 | 0.690 | 2.125 | 1.450 | 聚集型 |
6 | 19 | 2.000 | 1.564 | 2.222 | 0.222 | 1.111 | 0.111 | 1.222 | 8.998 | 聚集型 |
7 | 19 | 3.526 | 3.791 | 6.603 | 3.076 | 1.872 | 0.872 | 4.076 | 1.146 | 聚集型 |
8 | 19 | 6.105 | 5.557 | 10.163 | 4.057 | 1.665 | 0.665 | 5.057 | 1.505 | 聚集型 |
9 | 19 | 5.263 | 4.931 | 8.883 | 3.620 | 1.688 | 0.688 | 4.620 | 1.454 | 聚集型 |
10 | 19 | 3.263 | 2.535 | 4.233 | 0.970 | 1.297 | 0.297 | 1.970 | 3.366 | 聚集型 |
11 | 19 | 4.105 | 2.665 | 4.835 | 0.729 | 1.178 | 0.178 | 1.729 | 5.629 | 聚集型 |
12 | 19 | 5.158 | 3.338 | 6.318 | 1.160 | 1.225 | 0.225 | 2.160 | 4.447 | 聚集型 |
13 | 19 | 2.737 | 2.423 | 3.882 | 1.145 | 1.419 | 0.419 | 2.145 | 2.390 | 聚集型 |
14 | 19 | 3.263 | 3.088 | 5.186 | 1.923 | 1.589 | 0.589 | 2.923 | 1.697 | 聚集型 |
15 | 19 | 2.579 | 1.981 | 3.101 | 0.522 | 1.202 | 0.202 | 1.522 | 4.945 | 聚集型 |
不同虫态 Different insect states | 时间 Time | 模型 Model | 块金值C0 Nugget | 基态值 C0+C Sill | 变程a Range(km) | 空间结构比率 C0/(C0+C) Proportion of spatial structure (%) | 决定系数R2 Coefficient of determination | 空间格局 Spatial pattern |
---|---|---|---|---|---|---|---|---|
4月下旬 | 高斯型 | 36.5 | 267.7 | 0.000 88 | 13.634 665 67 | 0.894 | 聚集型 | |
5月上旬 | 高斯型 | 209 | 2 528 | 0.002 55 | 8.267 405 063 | 0.841 | 聚集型 | |
成虫 | 6月上旬 | 高斯型 | 10 | 21 120 | 0.001 37 | 0.047 348 485 | 0.967 | 聚集型 |
Adult | 7月中旬 | 高斯型 | 3 960 | 26 520 | 0.003 82 | 14.932 1267 | 0.543 | 聚集型 |
8月中旬 | 高斯型 | 10 | 13 670 | 0.000 54 | 0.073 152 89 | 0.712 | 聚集型 | |
8月下旬 | 线性 | 20 | 2 044 | 0.001 604 | 0.978 473 581 | 0.008 4 | 离散型 | |
5月中旬 | 球型 | 0.083 | 0.863 | 0.000 52 | 9.617 612 978 | 0.000 8 | 离散型 | |
6月上旬 | 球型 | 0.01 | 3.497 | 0.000 38 | 0.285 959 394 | 0.000 5 | 离散型 | |
幼虫 | 6月下旬 | 高斯型 | 0.49 | 31.97 | 0.001 65 | 1.532 686 894 | 0.618 | 聚集型 |
Larvae | 7月上旬 | 高斯型 | 10.1 | 221.1 | 0.015 7 | 4.568 068 747 | 0.943 | 聚集型 |
7月下旬 | 高斯型 | 0.1 | 175.2 | 0.001 53 | 0.057 077 626 | 0.948 | 聚集型 | |
8月上旬 | 高斯型 | 40.4 | 291.7 | 0.002 63 | 13.849 845 73 | 0.782 | 聚集型 |
Tab.4 Half variance function model parameters and spatial patterns of adult and larval populations of apple moth
不同虫态 Different insect states | 时间 Time | 模型 Model | 块金值C0 Nugget | 基态值 C0+C Sill | 变程a Range(km) | 空间结构比率 C0/(C0+C) Proportion of spatial structure (%) | 决定系数R2 Coefficient of determination | 空间格局 Spatial pattern |
---|---|---|---|---|---|---|---|---|
4月下旬 | 高斯型 | 36.5 | 267.7 | 0.000 88 | 13.634 665 67 | 0.894 | 聚集型 | |
5月上旬 | 高斯型 | 209 | 2 528 | 0.002 55 | 8.267 405 063 | 0.841 | 聚集型 | |
成虫 | 6月上旬 | 高斯型 | 10 | 21 120 | 0.001 37 | 0.047 348 485 | 0.967 | 聚集型 |
Adult | 7月中旬 | 高斯型 | 3 960 | 26 520 | 0.003 82 | 14.932 1267 | 0.543 | 聚集型 |
8月中旬 | 高斯型 | 10 | 13 670 | 0.000 54 | 0.073 152 89 | 0.712 | 聚集型 | |
8月下旬 | 线性 | 20 | 2 044 | 0.001 604 | 0.978 473 581 | 0.008 4 | 离散型 | |
5月中旬 | 球型 | 0.083 | 0.863 | 0.000 52 | 9.617 612 978 | 0.000 8 | 离散型 | |
6月上旬 | 球型 | 0.01 | 3.497 | 0.000 38 | 0.285 959 394 | 0.000 5 | 离散型 | |
幼虫 | 6月下旬 | 高斯型 | 0.49 | 31.97 | 0.001 65 | 1.532 686 894 | 0.618 | 聚集型 |
Larvae | 7月上旬 | 高斯型 | 10.1 | 221.1 | 0.015 7 | 4.568 068 747 | 0.943 | 聚集型 |
7月下旬 | 高斯型 | 0.1 | 175.2 | 0.001 53 | 0.057 077 626 | 0.948 | 聚集型 | |
8月上旬 | 高斯型 | 40.4 | 291.7 | 0.002 63 | 13.849 845 73 | 0.782 | 聚集型 |
允许误差D Allowable error | 平均密度Mean density | |||||||
---|---|---|---|---|---|---|---|---|
3 | 5 | 7 | 10 | 15 | 20 | 25 | 30 | |
0.05 | 3 432 | 2 641 | 2 210 | 1 966 | 1 776 | 1 681 | 1 624 | 1 586 |
0.10 | 349 | 350 | 553 | 492 | 444 | 420 | 406 | 397 |
0.20 | 88 | 159 | 87 | 123 | 111 | 105 | 102 | 99 |
0.30 | 92 | 70 | 61 | 55 | 49 | 47 | 45 | 44 |
Tab.5 Theoretical sampling numbers of adult apple moth under different population densities
允许误差D Allowable error | 平均密度Mean density | |||||||
---|---|---|---|---|---|---|---|---|
3 | 5 | 7 | 10 | 15 | 20 | 25 | 30 | |
0.05 | 3 432 | 2 641 | 2 210 | 1 966 | 1 776 | 1 681 | 1 624 | 1 586 |
0.10 | 349 | 350 | 553 | 492 | 444 | 420 | 406 | 397 |
0.20 | 88 | 159 | 87 | 123 | 111 | 105 | 102 | 99 |
0.30 | 92 | 70 | 61 | 55 | 49 | 47 | 45 | 44 |
允许误差D Allowable error | 平均密度Mean density | |||||||||
---|---|---|---|---|---|---|---|---|---|---|
1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | |
0.05 | 1 738 | 1 361 | 1 235 | 1 172 | 1 134 | 1 109 | 1 091 | 1 078 | 1 067 | 1 059 |
0.10 | 435 | 340 | 309 | 293 | 284 | 277 | 273 | 269 | 267 | 265 |
0.20 | 109 | 85 | 77 | 73 | 71 | 69 | 68 | 67 | 67 | 66 |
0.30 | 48 | 38 | 34 | 33 | 32 | 31 | 30 | 30 | 30 | 29 |
Tab.6 Theoretical sampling numbers of apple moth larvae under different population densities
允许误差D Allowable error | 平均密度Mean density | |||||||||
---|---|---|---|---|---|---|---|---|---|---|
1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | |
0.05 | 1 738 | 1 361 | 1 235 | 1 172 | 1 134 | 1 109 | 1 091 | 1 078 | 1 067 | 1 059 |
0.10 | 435 | 340 | 309 | 293 | 284 | 277 | 273 | 269 | 267 | 265 |
0.20 | 109 | 85 | 77 | 73 | 71 | 69 | 68 | 67 | 67 | 66 |
0.30 | 48 | 38 | 34 | 33 | 32 | 31 | 30 | 30 | 30 | 29 |
项目 Items | 设置诱捕器The number of traps(n) | |||||||||
---|---|---|---|---|---|---|---|---|---|---|
5 | 10 | 15 | 20 | 25 | 30 | 35 | 40 | 45 | 50 | |
下限T0(n) | 13 | 39 | 66 | 92 | 118 | 144 | 170 | 197 | 223 | 249 |
上限T1(n) | 39 | 66 | 92 | 118 | 144 | 171 | 197 | 223 | 249 | 275 |
Tab.7 Iwao sequential sampling sequence of adult apple moth
项目 Items | 设置诱捕器The number of traps(n) | |||||||||
---|---|---|---|---|---|---|---|---|---|---|
5 | 10 | 15 | 20 | 25 | 30 | 35 | 40 | 45 | 50 | |
下限T0(n) | 13 | 39 | 66 | 92 | 118 | 144 | 170 | 197 | 223 | 249 |
上限T1(n) | 39 | 66 | 92 | 118 | 144 | 171 | 197 | 223 | 249 | 275 |
项目 Items | 调查果数Survey number(n) | |||||||||
---|---|---|---|---|---|---|---|---|---|---|
100 | 200 | 300 | 400 | 500 | 600 | 700 | 800 | 900 | 1000 | |
下限T0(n) | 1 | 4 | 6 | 9 | 11 | 14 | 16 | 18 | 21 | 23 |
上限T1(n) | 4 | 6 | 9 | 11 | 14 | 16 | 19 | 21 | 24 | 26 |
Tab.8 Iwao sequential sampling sequence of apple moth larvae
项目 Items | 调查果数Survey number(n) | |||||||||
---|---|---|---|---|---|---|---|---|---|---|
100 | 200 | 300 | 400 | 500 | 600 | 700 | 800 | 900 | 1000 | |
下限T0(n) | 1 | 4 | 6 | 9 | 11 | 14 | 16 | 18 | 21 | 23 |
上限T1(n) | 4 | 6 | 9 | 11 | 14 | 16 | 19 | 21 | 24 | 26 |
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